When puffins are threatened by a predator, such as a fox or an eagle, they can defend themselves by squirting a foul-smelling oil from a gland near their tail. The oil is effective at repelling predators, but it can also be harmful if it gets in the eyes of a human or another animal.
Biting
Puffins also have a sharp beak that they can use to defend themselves. If they are attacked by a predator, they will often try to bite the predator's nose or eyes. This can be an effective way to deter a predator, but it can also be dangerous for the puffin if the predator is able to fight back.
Kicking
Puffins can also use their feet to defend themselves. They can kick their predators with their sharp claws, which can cause pain and injury. This is often a last resort for puffins, but it can be an effective way to deter a predator if they are able to land a solid kick.